Biology

I. Skhirtladze

The List of Bees of Caucasus

(Hymenoptera: Colletidae, Melittidae)

The Stdly of bees of Caucasus began in the first of the XIX century [1: 110-133; 2: 161-165; 3: 181-202; 4: 268-273; 5: 61-64]. But planned investigation of bees of Caucasus began only in the second half of the XX century [6: 191-205; 7: 270-289; 8: 134-153; 9: 337-348; 10: 3-17; 11: 3-13; 12: 112-121; 13: 92-187; 14: 60-98; 15: 318-350; 16: 520; 17: 133-154; 18: 16-65; 19: 189; 20: 129-189]. In this scientific work the list of bees’ families Colletidae and Melittidae Caucasus is represented compatibly fully for the first time.

The Colletidae family was determined to incldle 61 species of 2 subfamily, 2 genera and the Melittidae family was determined to incldle 14 species, of 3 subfamily and 3 genera.

Ordo - Hymenoptera*
Super family - Apoidea, Ashmead, 1899
I. Family - Colletidae Bingham, 1897
I. Subfamily - Colletinae Bönner, 1919
Genus - Colletes Latreille, 1802
  1. Colletes carinatus Radoszkowski, 1891
    Distribution: Eastern Georgia: Borjomi,*
    Azerbaijan: Kirovobad, Zangelan v. Vezhnali [7; 9]
    Number: Rare
  2. C. daviesanus Smith, 1846
    Distribution: Eastern Georgia: Tsodoreti (Tbilisi), Lagodekhi – national park, Shroma (Lagodekhi), Matsim-chai (Lagodekhi), Rachisubani (Lagodekhi), Khizabavra (Lagodekhi), Kaspi [21: 3-48], Tvalivi (Dusheti) [8].
    Western Georgia: Glola (Oni) [3], Bichvinta-national park [22: 2-23],
    Azerbaijan: Kuba [16].
    Armenia: Echmiadzin, Leninakan [23: 3-69].
    Number: Ordinary
  3. C. hylaeiformis Eversmann, 1852
    Distribution: Eastern Georgia: Tbilisi.**
    Azerbaijan: Mingechaur,*** Lerik [17]
    Number: Not numerous
  4. C. marginatus Smith, 1846
    Distribution: Eastern Georgia: Tsodorethi (Tbilisi), Lagodekhi – national park; Shroma (Lagodekhi), Kaspi [21], Batsara-Babaneuri – national park.**** Liakhvi – national park [11].
    Western Georgia: Mulakhi (Mestia) [12]; Liakhvi (Sauth Osetia) – national park [11], Glola (Oni) [3; 18].
    Azerbaijan: Kyurdamir – (steppe Shirvani); steppe Mili; Kizilagakh – national park, Dzhulpha (Nakhichevan).****
    North Caucasus: Lake Adji (Daghestan) [24: 127-144].
    Number: Ordinary
  5. C. mlokosiewiczi Radoszkowski, 1891
    Distribution: Eastern Georgia: Tbilisi*, Lagodekhi – national park*, Batsara-Babaneuri – national park (Akhmeta)****
    Number: Not numerous
  6. C. pallescens Noskiewicz, 1936
    Distribution: Eastern Georgia: Lagodekhi – national park, Shroma (Lagodekhi), Rachisubani (Lagodekhi) [9], ravine Kumuro – Vashlovani – national park [25:145-160];
    Western Georgia: Pharakheti (Oni) [3], Mtiskalta [18].
    Azerbaijan: Ordubad, Kelbadjar, Zhdanovsk (steppe Mili), Nakhichevan, ravine Pragachai [26: 115 145].
    North Caucasus: Lake Adji (Daghestan) [24].
    Number: Ordinary
  7. C. similis Schenck, 1853
    Distribution: Eastern Georgia: Borjomi**, Lagodekni** - national park;
    Western Georgia: Chkaduashi (Zugdidi), Akhalsopheli (Zugdidi), Latsurtsume (Chkhorotsqu [1], Gagra – Bichvinta – national park [22].
    Number: Rare
  8. C. nasutus Smith, 1859
    Distribution: Eastern Georgia: Tbilisi***, Atsquri [23];
    Azerbaijan: Shemakha****;
    North Caucasus: ravine Djariakhi, village Armkhi (Checheno-Ingusheti)****.
    Number: Rare
  9. C. spectabilis F.Morawitz, 1868
    Distribution: Eastern Georgia: Tbilisi***;
    North Caucasus: Derbent (Daghestan) [20; 24].
    Number: Rare
  10. C. cunicularius Linnaeus, 1758
    Distribution: Eastern Georgia: Tbilisi***;
    Western Georgia: Sairme (maiakovsk), Lake Paliastomi (Poti) [22];
    Azerbaijan: Mingechaur, Saatly (steppe Mili)****, Baku [20], Zuvand, Gosmolian [17], Kirovobad [16];
    Armenia: Khasro – national park****.
    Number: Not numerous
  11. C. tuberculatus F.Morawitz, 1894
    Distribution: Eastern Georgia: Tbilisi***;
    Azerbaijan: v. Bichenek, Shakhbuz [27: 202-210; 21].
    Number: Rare
  12. C. fodiens Geoffroy, 1762
    Distribution: Eastern Georgia: Tbilisi***, mountain Mamadaviti****, ravine Katsalkhevi (Dusheti) [6];
    Azerbaijan: Kyurdamir, Kerrar [27];
    North Caucasus: Dombai (Karachaevo-Cherkesk), National park – Caucasus****.
    Number: Not numerous
  13. C. lebedewi Noskeiwicz, 1936
    Distribution: Eastern Georgia: Tbilisi,***
    Azerbaijan: Kirovabad,**** Kutkashen [27].
    Number: Rare
  14. C. eous Morice, 1904
    Distribution: Eastern Georgia: Tbilisi.***
    Azerbaijan: Zangelan, v. Vezhnali [27].
    Number: Rare
  15. C. mountainous F. Morawitz, 1876
    Distribution: Eastern Georgia: Phasanauri [28].
    Number: Rare
  16. C. transitorius Naskiewicz, 1936
    Distribution: Azerbaijan: Nakhichevan, v. Buzgav [27];
    North Caucasus: Kabardino-Balkaria-heightmauntain national park.****
    Number: Not numerous.
  17. C. succinctus (Linnaeus, 1758)
    Distribution: Azerbaijan: Kilyaze, Tazakend [27];
    North Caucasus: Curusch (Daghestan) [20].
    Number: Rare
  18. C. squamosulus Noskiewicz, 1936
    Distribution: North Caucasus: Karachaevo-Cherkesk-pass Klukhori, National park – Caucasus.****
    Number: Rare
  19. C. caspicus F.Mor, 1873
    Distribution: Azerbaijan: Kirovabad.****
    North Caucasus: Derbent (Daghestan) [20].
    Number: Rare
  20. C. floralis Eversm, 1852
    Distribution: Western Georgia: Khulo (Adzharia);****
    Azerbaijan: Kirovobad [5];
    North Caucasus: Tsei-glacier Skazka (North Osetia)****.
    Number: Rare
  21. C. maidli Noskiewicz, 1936
    Distribution: Western Georgia: Maiakovsk, Adjameti – national park [22].
    Number: Rare
  22. C. punctatus Mocsary, 1877
    Distribution: Azerbaijan: v. Laza, Shakhdag,**** Zangelan, v. Vezhnali [27].
    Number: Rare
  23. C. graeffei Alfken, 1900
    Distribution: Azerbaijan; Kuba [27].
    Number: Rare
  24. C. cretaceus F. Mor., 1876
    Distribution: Armenia: Erevan.***
    Number: Rare
  25. C. comatus Noskiewicz, 1936
    Distribution: Armenia: Vedi, Sarai-bulag.****
    Number: Rare
II. Subfamily: Prosopinae Börner, 1919
=Hylaenae, Michener, 1944
Genus Prosopis Fabricius, 1804
  1. Prosopis cornuta Smith, 1842
    Distribution: Eastern Georgia: Tbilisi [21], Lagodekhi – national park, Sartichala [13; 28], Diklo (Akhmeta) [10];
    Western Georgia: Pitsunda – national park [22];
    Azerbaijan: Sheki [13; 28], Mardakert, v. Leninaval [27];
    North Caucasus: Jargil (Daghestan), Gilari (Daghestan) [24].
    Number: not numerous
  2. P. punctulatissima Smith, 1842
    Distribution: Eastern Georgia: Borjomi, Tbilisi [13];
    Azerbaijan: Ordubad [27].
    Number: Rare
  3. P. bifasciata jurine, 1807
    Distribution: Eastern Georgia: Tbilisi;***
    Azerbaijan: Kirovobad, Nakhichevan [27];
    Norht Caucasus: Khosta (Krasnodar).****
    Number: Rare
  4. P. variegata Fabricius, 1798
    Distribution: Eastern Georgia: Vashlovani – national park [25], Omalo (Akhmeta) [10];
    Azerbaijan: Shakhbuz, v. Kiu-kiu, Dzhulfa, v. Bilav,**** Girkani – national park,**** Bacu, Kirovobad, Akstapha [13], Mardakert, v. Leninaval [27];
    Armenia: Erevan [13];
    North Caucasus: Derbent (Daghestan) [19], Akhti (Daghestan) [1-2; 9], Jargil (Daghestan), Gilari (Daghestan) [24], river Kuban (Stavropol)****.
    Number: Ordinary
  5. P. gibba Saunders, 1850
    Distribution: Eastern Georgia: Lagodekhi – national park, Tbilisi [1; 7];
    Azerbaijan: Mardakert, v. Leninaval, Kirovobad [27].
    Number: Rare
  6. P. confusa Nylander, 1852
    Distribution: Eastern Georgia: Borjomi, Abastumani [13; 23; 28];
    Western Georgia: Liakhvi – national park [11];
    Azerbaijan: Ordubad**** Kuba [16];
    North Caucasus: Blata (Vladikavkaz).****
    Number: Not numerous
  7. P. pratensis Geoffroy, 1785 = signata Fabricius, 1804
    Distribution: Eastern Georgia: Tbilisi, Borjomi [2];
    Azerbaijan: Ordubad [27], Kirovobad [13; 23], Akstapha [23].
    Number: Rare
  8. P. brevicornis Nylander, 1852
    Distribution: Eastern Georgia: Lagodekhi – national park, Shroma (Lagodekhi), Khizabavra (Lagodekhi), Tbilisi, Kodjori [13], Borjomi [28];
    Western Georgia: Akhalsopheli (Zugdidi), Garakha (Chkhorotsku) [1], Besleti (Sokhumi), Kelasuri (Sokhumi) [23], Kutaisi [21];
    Azerbaijan: Mardakert, v. Leninaval [27], Kelbadjar.****
    Number: Ordinary
  9. P. hungarica Alfken, 1905
    = affinis Mor. 1876
    = Morowitz D.T. 1896
    Distribution: Eastern Georgia: Borjomi [13; 23].
    Number: Rare
  10. P. pictipes Nylandez, 1852
    Distribution: Eastern Georgia: Borjomi, Lagodekhi, Tbilisi [13; 23].
    Number: Rare
  11. P. minuta Fabricius, 1798 = Sinuata
    Distribution: Eastern Georgia: Tbilisi;***
    Western Georgia: Glola (Oni) [3];
    Azerbaijan: Mardakhert v. Talish [27].
    Number: Rare
  12. P. lineolata Schenck, 1859
    Distribution: Eastern Georgia: Tbilisi, Borjomi, Lagodekhi, Kodjorik, Mtskheta [13; 23];
    Azerbaijan: Dzhulpha (Nakhichevan), ravine Paragachai, v. Bilav,**** Mardakert, v. Leninaval [27].
    Number: Not numerous
  13. P. angustata Schenck, 1859
    Distribution: Eastern Georgia: Tbilisi, Lagodekhi – national park, Kodjori [28];
    Azerbaijan: Kelbadjar,**** Kazakh, v. Djapharli [27].
    Number: Rare
  14. P. bisinuata Förster, 1871
    Distribution: Eastern Georgia: Lagodekhi – national park [21];
    Western Georgia: Besleti (Sokhumi), Gumista (Sokhumi), Gagra, Lake Ritsa, Gdlauta [23];
    Azerbaijan: Kelbadjar, Bilav (Nakhichevan),**** Mardakert, v. Leninaval, Kazakh, v. Djapharli [27];
    North Caucasus: Makhachkala (Daghestan) [24].
    Number: Orginary.
  15. P. communis Nylander, 1852
    Distribution: Eastern Georgia: Lagodekhi – national park Shroma (Lagodekhi) [9], Vashlovani – national park [25], Tsalka [7; 18: 16-65], Batsara-Babaneuri – national park,**** Borjomi [23; 29: 263-267], Akhaltsikhe [6];
    Western Georgia: Akhalsopheli (Zugdidi) [1], Utsera (Oni), Seva (Oni) [3; 18], Mulakhi (Mestia) [12], Liakhvi – national park (South Osetia) [11];
    Azerbaijan: v. Nius-nius (Nakhichevan), Kelbadjar,**** Mardakert, v. Leninaval, Zangelan v. Vezhnali, Khanlar, Lake Gek-gel [27], Kirovobad [5];
    North Caucasus: Pyatigorsk [23], Tuaphse-Khosta (Krasnodar territory), ****.
    Number: Numerous
  16. P. nigrita Fabricius, 1798 = propinquus Nylander, 1852
    Distribution: Eastern Georgia: Kasbegi [13; 28], Roshka pass (Dusheti) [5];
    Azerbaijan: Shakhbuz pass Bichenek, ravine Paragachai, v. Bilav (Nakhichevan).****
    Number: Rare
  17. P. difformis Eversmann, 1852 = subfasciata Schenk, 1874
    Distribution: Eastern Georgia: Phasanauri [13; 23];
    Western Georgia: Ushguli (Mestia) [12];
    Azerbaijan: Zangelan, v. Vezhnali [27].
    Number: Not numerous
  18. P. annulata Linne, 1758 = borealis Nyl
    Distribution: Eastern Georgia: Lagodekhi – national park, Rachisubani (Lagodekhi), Matsim-chai (Lagodekhi), Gurgeniani (Lagodekhi) [9], Vashlovani – national park, ravine Kumuzo (Vashlovani) [25], Kasris tskali (steppe Eldari) [14], Batsara-Babaneuri – national park****.
    Western Georgia: Kutaisi [7], Lidzava (Gagra) [22], Kintrishi – national park [2];
    Azerbaijan: Mardakert v. Leninaval [2], Ordubad, Kobistan, ravine Paragachai (Nakhichevan), between Ordubod and Megri;****
    Armenia: Megri.****
    Number: Numerous
  19. P. annularis Kirby, 1802 = dilitata Kirby, 1802
    Distribution: Eastern Georgia: Lagodekhi, Borjomi [28], ravine Datvisi (Dusheti) [8];
    Western Georgia: Chkaduashi (Zugdidi), Letsurtsume (Chkorotsku) [1; 22];
    Azerbaijan: Mardakert, v. Leninaval [27].
    Number: Not numerous
  20. P. euryscapa Förster, 1871 = rimosa
    Distribution: Eastern Georgia: Tbilisi;***
    Azerbaijan: Mardakert, v. Leninaval [27].
    Number: Rare
  21. P. hyalinata Smith, 1842
    Distribution: Eastern Georgia: Borjomi, Qasbegi [13];
    Azerbaijan: Kelbadjar, health Istisu; Mardakert, v. Leninaval [27];
    Armenia: Yerevan [13].
    Number: Not numerous
  22. P. ibex F. Morawitz, 1878
    Distribution: Eastern Georgia: Gdlauri, Qasbegi [13].
    Number: Rare
  23. P. armillatus Nylander, 1852
    Distribution: Eastern Georgia: Borjomi [23];
    Armenia: Erivan [23].
    Number: Rare
  24. P. rugicollis F. Mor, 1873
    Distribution: Azerbaijan: Kirovobad;****
    North Caucasus: Derbent (Daghestan) [13; 20].
    Number: Rare
  25. P. punctata Brulle, 1832
    Distribution: Azerbaijan: v. Laza, Shakh-dag.****
    Number: Rare
  26. P. scutellaris F. Mor. 1873
    Distribution: Azerbaijan: Baku [20].
    Number: Rare
  27. P. breviventris F. Först, 1871
    Distribution: North Caucasus: Derbent (Daghestan) [13; 20]
    Number: Rare
  28. P. breviceps F. Mor., 1876
    Distribution: Eastern Georgia:
    Azerbaijan: Ordubad (Nakhichevan),**** Mardakert v. Leninaval [27].
    Number: Rare
  29. P. obscuratus Schenck, 1859
    Distribution: Eastern Georgia: Borjomi [23].
    Number: Rare
  30. P. clippearis Schenck, 1853
    Distribution: Kirovobad.****
    Number: Rare
  31. P. rinki Gorski, 1852
    Distribution: Azerbaijan: Mir-Basher, v. Chile [27].
    Number: Rare
  32. P. gracilicornis F. Mor., 1868
    Distribution: Azerbaijan: Ordubad, ravine v. Bilav.****
    Number: Rare
  33. P. duckei Alfken, 1904
    Distribution: Azerbaijan: Shakhbuz, v. Bichenek, **** Kelbadjar, riv. Terter, Mardakert, v. Leninaval [27].
    Number: Not numerous
  34. P. imparilis Förster, 1871
    Distribution: Azerbaijan: Mardakert, v. Leninaval, v. Chile [27].
    Number: Rare
  35. P. styriacus Förster, 1871
    Distribution: Azerbaijan, Mardakert, v. Leninaval, Ordubad [27].
    Number: Rare
  36. P. moricei luteifrons Strand, 1913
    Distribution: Azerbaijan: Mardukert v. Leninaval [27].
    Number: Rare
II. family – Melittidae
I. Subfamily – Melittinae Schenck, 1859
Genus – Melitta Kirby, 1802
  1. Melitta haemorrhoidalis Fabricius, 1775
    Distribution: Eastern Georgia: Borjomi,** Abastuman;**
    Azerbaijan: Khanlar, Lake Gei-gel [5];
    North Caucasus: Curusch (Daghestan) [20]
    Number: Rare
  2. M. leporina Panzer, 1799
    Distribution: Eastern Georgia: Borjomi;****
    Azerbaijan: Gosmolian (Lenkoran), Gilidara, Zuvand [17], Mardakert, v. Leninaval [5];
    Armenia: Leninakan [23].
    Number: Not numerous
  3. M. dimidiate Mor., 1876
    Distribution: Azerbaijan: ravine Paragachai (Nakhichevan) v. Bilav,**** Mastara [23], Mardakert, v. Talish [5].
    Number: Rare
  4. M. tricincta Kby, 1851 = melanura Nyl, 1852
    Distribution: Azerbaijan: Mardakert, v. Leninaval [5].
    Number: Rare
II. Subfamily Dasypodinae
Genus Dasypoda Latreille, 1802
  1. Dasypoda argentata Panzer, 1809
    Distribution: Eastern Georgia: Shatili (Dusheti) [8], Abastuman [9];
    Armenia: Evevan [9].
    Number: Rare
  2. D. plumipes Panzer, 1809
    Distribution: Eastern Georgia: Tbilisi;**
    Western Georgia: Kintrishi – national park [2];
    Azerbaijan: Akstapha [9], v. Balachali [5];
    North Caucasus: Derbent (Daghestan) [13; 20].
    Number: Not numerous
  3. D. vasnaga Rossi, 1790
    Distribution: North Caucasus: Derbent (Daghestan) [9; 153; 15].
    Number: Rare
  4. D. schletereri Friese, 1885
    Distribution: North Caucasus: Derbent (Daghestan) [15].
    Number: Rare
  5. D. spinigcra Kohl, 1905
    Distribution: Azerbaijan: Shakhbuz, v. Bichenek [5].
    Number: Rare
  6. D. mlokosievitzi Radoszkowski, 1891
    Distribution: Azerbaijan: Shakhbuz, v. Kiu-Kiu.****
    Armenia: Erevan [15].
    Number: Rare
  7. D. subinterrupta Brulle, 1832
    Distribution: North Caucasus: Derbent (Daghestan) [20].
    Number: Rare
III. Subfamily Macropodinae Michn, 1944
Genus Macropis Klug, 1809
  1. Macropis Fulvipes Fabricius, 1804
    Distribution: Western Georgia: Orule (Zugdidi) [1].
    Number: Rare
  2. M. labiata Fabriacius, 1804
    Distribution: Eastern Georgia: Lagodekhi – national park [21; 28];
    Western Georgia: v. Orule (Zugdidi) [1].
    Number: Rare
  3. M. nigrita Hedicke, 1933
    Distribution: Armenia: Erevan [15];
    North Caucasus: Territory Krasnodar, Station Lazarevskaia [15].
    Number: Rare

Transcription

*
Collection institute Zoology St.-Petersburg
**
Collection Zoo Museum Moscow
***
Collection institute Zoology Kiev
****
Collection Museum Georgia

 

S. Janashia Georgian State Museum
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